Making Smart Assumptions about Prairie Management

Some people say it’s dangerous to make assumptions.  I disagree.  In fact, assumptions are both necessary and empowering.  Land managers make assumptions all the time.  If we didn’t, we’d never get anything done.

Assumptions are only dangerous when they are either unrecognized or untested.  For example, it’s reasonable for me to assume that my car’s engine has an adequate amount of oil in it, but it would be irresponsible not to check the level now and then to be sure.  Without the assumption that I still had oil, I’d probably stop to check my oil every mile or so and I’d never get anywhere.  In order to move forward, I have to make reasonable assumptions, including that my engine hasn’t lost all of its oil since the last time I checked it.

As land managers, we have to take a similar approach.  Much of the time, we assume that species and ecological systems are reacting predictably and positively to our management, but we also do spot checks to reassure ourselves.  Often those evaluations involve nothing more than a walk through a prairie to see how things are looking, but in some cases might conduct more intensive data collection.

Sometimes rigorous data collection is used to test assumptions, but other times the process can be much more informal.

Sometimes rigorous data collection is used to test assumptions, but other times the process can be much more informal.

Land managers also make broader assumptions about how restoration or management projects will contribute to conservation objectives.  As we plan projects, we make educated guesses that help us design our work effectively.  Then we implement the project and see what happens.  If we didn’t make assumptions, we’d be paralyzed by indecision and never get anything done.

It is critically important, however to recognize what assumptions we’re making, and to test them when we have the chance.  Here are several examples of assumptions we make in our Platte River Prairies management, and some of the ways we’re testing them.

Assumption #1.  Prairie plants can survive periodic intensive grazing.

Grazing is an important part of our management.  Most commonly, we employ variations of patch-burn grazing, in which a portion of a prairie is grazed pretty intensively for most of one growing season and then allowed to recover for a couple years before it’s intensively grazed again.  We use grazing to manipulate plant competition, especially by suppressing the vigor of dominant grasses to produce more plant diversity.  It also is our primary tool for creating heterogeneous habitat structure, including important habitat conditions (such as short grass/tall forbs) that are difficult or impossible to create without grazing.

Selective grazing by cattle can have positive impacts on both plant diversity and wildlife habitat.  These cattle at Konza Prairie in Kansas are grazing on primarily grass and leaving plants such as leadplant and purple prairie clover ungrazed.

Selective grazing by cattle can have positive impacts on both plant diversity and wildlife habitat. These cattle at Konza Prairie in Kansas are grazing on primarily grass and leaving plants such as leadplant and purple prairie clover ungrazed.

Our data show that overall plant diversity is thriving under our management, and it’s easy to see the variety of habitat conditions we create each year.  However, we’re making the assumption that we’re not losing any plant species due to periodic intensive grazing.  It’s an informed assumption, based on experience and our understanding of history, including the kind of fire/grazing interactions that happened in these prairies over the last several thousand years.  Regardless, it’s an assumption, and one we need to test.

We collect annual data from some prairies and less frequent data from others that allow us to track individual plant species over time.  So far, we’ve not seen any indication of plant species that are disappearing under our management.  Even if we weren’t rigorously collecting data, we could still test our assumption by simply tracking the population size of species most likely to be impacted by grazing.  We could use techniques such as photopoints or walking transects, or we could just mark and watch individuals or patches of plants over time.

Data

An example of the kind of data that helps us evaluate the impacts of repetitive grazing on plant species.  Data from one of our restored prairies shows plant species that have maintained populations through 13 years of patch-burn grazing.  The data represent the frequency of occurrence for each species within approximately 100 plots (1 square meter plots).  From top to bottom, the common names of the species shown are Stiff Sunflower, Maximilian Sunflower, Illinois Bundleflower, Purple Prairie Clover, and White Prairie Clover.

Assumption #2.  Some exotic/invasive species are not harmful enough to warrant eradication efforts.

We have more than enough invasive species to deal with on our sites, so we have to be selective about which to spend most of our time on.  We set priorities based on experience, and focus most on those species we think have the greatest potential to harm plant diversity or habitat quality.  However, we recognize that our assumptions about impacts could be wrong, so we test them – both through general observation and data collection.  I’ve written before about data we’ve collected on both Kentucky bluegrass and sweetclover impacts, for example.

Assumption #3.  Restoring cropland adjacent to a small prairie will increase its conservation value.

The prairie restoration work we do is not focused on re-creating historic landscapes, but on trying to decrease the impacts of habitat fragmentation.  We assume that adding diverse prairie plantings around and between isolated prairie fragments will increase population size and connectivity for plants, insects, mammals, birds, reptiles, and more.  Bigger and more interconnected populations should be more viable than smaller and isolated populations.

Our assumption seems reasonable, but it’s expensive to harvest and plant more than 200 plant species in a crop field, so we need to see if we’re actually achieving our objective.  More importantly, we need to be able to show policy makers that this kind of strategy produces substantial ecological impacts.  Unfortunately, this kind of assumption is logistically difficult to test.

We have a long way to go, but we’re starting to look at whether various species living in our unplowed prairies are also found in adjacent restored prairies.  If those species aren’t using the new habitat, our strategy isn’t helping them.  If they are, that’s good to know – though there are still more assumptions to test (e.g., do those new habitats facilitate successful breeding and/or migration and colonization?).  So far, some preliminary investigations indicate that most ant and bee species appear to use restored habitats, and we’re now looking at small mammals and grasshoppers as well.

ENPO130528_D023

Bees seem to readily use the restored prairie habitats we create around and between formerly-isolated prairies. However, we are still evaluating bees and many other species to see how effective our restoration work has been.

 

Assumption #4.  We can maintain healthy populations of all prairie species through our “shifting mosaic” approach to wildlife habitat management.

This is a big one, and is very difficult to test.  We assume that by creating a variety of habitat conditions each year – including tall/dense, short/sparse, and mixed-height vegetation – all of the species in our prairies (insects, mammals, birds, plants, etc.) can find what they need each year.  On top of that, we’re assuming that as we shift the location of habitat conditions between years, species can either move to appropriate habitat or hunker down until better conditions cycle back through.

As with our assumption about plants and grazing, historical context applies.  Prairie species evolved in grasslands that were subjected to fire, grazing, and drought, and preferred habitat conditions for any particular species would have shifted around the landscape from year to year.  However, much is different today, including the size and fragmentation of grasslands, the presence of invasive species, and much more.  Are today’s species able to survive significant variations in habitat conditions from year to year?  Can a species that needs thatchy cover successfully find more of that habitat after a fire burns through its current location?  If so, how far can it travel, and through what kinds of habitats?

Red

How easily can a species such as the red-sided garter snake find new habitat after a fire or other management treatment makes its current habitat unsuitable?

We haven’t gotten as far in testing this assumption as we have with some others, but we think a lot about it.  We’ve been gleaning information on animal movement from the scientific literature, and will meet with university scientists next month to discuss potential collaborative research on this topic.  Most importantly, we recognize that we are making some big assumptions about our management strategies, and we keep those assumptions in mind as we make our annual plans.  For example, we try to think about factors such as travel distance between similar habitat types (tall/dense or short/sparse habitats, for example) and we try to leave unburned refuges within large burn units.  Hopefully, we’ll get more guidance soon, but in the meantime, we’re moving forward the best we can.

Just as I watch for signs that my car’s engine might be getting low on oil as I drive, I also watch for signs that our land management strategies are working as we want them to.  (More on that in an upcoming post.)  Recognizing the assumptions we’re making is a critical piece of successful management, but testing those assumptions is just as important.  Assumption testing doesn’t have to involve intensive data collection; it can be as simple as making some annual notes about whether or not a particular patch of wildflowers is still there, or keeping track of how invasive species respond to various management treatments.  If we know what we’re uncertain of, we’ll be more thoughtful about management decisions and more observant of their impacts.

What assumptions are you making?  Are you working to test them?

…oh, and don’t forget to check your oil now and then.

 

 

 

So Similar, Yet So Different

It’s wrong to assume that successful restoration or management tactics from one prairie will work in another.  Instead, every prairie has its own “personality” and responds accordingly.  The key to success is experimentation and adaptive management.

Bill Kleiman is one of my favorite people.  We have much in common: a love of prairies and restoration, a drive to learn from our mistakes and share what we learn with others, and a strong belief in the importance of conservation.  We’ve both worked for The Nature Conservancy for a long time (he’s got a couple years on me) and have been co-leading the Grassland Restoration Network for the last several years.  He’s also a great guy and a good friend.

Bill Kleiman (white hat) leads a tour of a restored prairie at The Nature Conservancy's Nachusa Grassland.

Bill Kleiman (light-colored hat) leads a tour of a restored prairie at The Nature Conservancy’s Nachusa Grasslands.  2014 Grassland Restoration Network workshop.

However, despite the fact that Bill and I are friends and have a lot in common, there are some big differences between us as well.  Bill is much more patient than I am, and better at the social niceties needed to build close relationships with neighbors and volunteers.  I tend to say what I think – sometimes inappropriately.  Bill is not shy about expressing his opinion, but does it less frequently, and usually with kindness and self-deprecation.

Bill and I both manage grasslands for The Nature Conservancy, but just as there are differences between us as people, there are also some stark differences between our sites and the approaches we take toward prairie restoration and management.  Bill’s site, TNC’s Nachusa Grasslands, is located in rolling hills about two hours west of Chicago, Illinois.  My Platte River Prairies are on mostly flat alluvial (river-formed) soils in south-central Nebraska.  As a result, the soils, topography and climate vary greatly between the two sites.  Moreover, our sites each have unique land use histories, invasive species legacies, and social and cultural contexts.

Bill again, talking to a tour group - with the Nachusa Grasslands in the background.

Bill again, talking to the same tour group, with Nachusa Grasslands’ undulating topography in the background.

I was thinking about all of this last month as our Platte River Prairies crew traveled to Nachusa Grasslands where Bill and his team were hosting this year’s annual Grassland Restoration Network workshop.  It was fascinating to compare the land management and restoration strategies we each use, especially knowing that both of us have diligently tested and refined our methods to meet the individual challenges of our respective sites.  Below are some of the similarities and differences between our approaches.

1. Seed Mixtures and Seeding Rates

Both Nachusa Grasslands and the Platte River Prairies have been actively restoring cropland to high-diversity prairie habitats.  At both sites we broadcast our seed (as opposed to drilling it).  The seed is broadcast either by hand or with a drop spreader – a fertilizer spreader that drops seeds onto the ground.  In fact, broadcast seeding is the technique of choice for the vast majority of sites that participate in the Grassland Restoration Network.  (You can learn more about fairly universal strategies and tactics in the “Lessons from the Grassland Restoration Network” document several of us put together.)

However, while we both broadcast seeds, Bill has found that successful prairie plantings at Nachusa require much heavier seeding rates (around 50 bulk pounds of seed per acre) than we use along the Platte River (8-10 bulk pounds).  Bill’s seed mixes include lots of seed from wildflowers, sedges, and “subdominant” grasses such as little bluestem, prairie dropseed, and sideoats grama, but almost no seed from more dominant grasses such as big bluestem and indiangrass.  In our Platte River Prairies seed mixtures, dominant grasses make up about half of the weight of the mixture.

James Trager (Shaw Nature Reserve) and Nelson Winkel (TNC Platte River Prairies) look over a restored prairie at Nachusa Grasslands.  This prairie was relatively unique in that it had a fair amount of indiangrass in it.  Most of the seed mixtures have none, or very little, seed from big grasses such as indiangrass and big bluestem.

James Trager (Shaw Nature Reserve) and Nelson Winkel (TNC Platte River Prairies) look over a restored prairie at Nachusa Grasslands. This prairie was relatively unique in that it had a fair amount of indiangrass in it. Most of the seed mixtures have none, or very little, seed from big grasses such as indiangrass and big bluestem.

Both Bill and I have experimented with many variations of these seed mixtures and have settled on these broad recipes as appropriate for our respective sites.  Though we do things differently, we both end up with very diverse prairies that meet our objectives.  When Bill uses lighter seeding rates, his new prairies get swamped out by invasive species before native plants become well established.  He’s also found that adding dominant grasses to the initial seed mix leads to plant communities that become overly grassy and not very diverse.  In contrast, using lighter seeding rates on the Platte allows us to plant more acres per year with the same seed harvest effort, and while it takes longer for our plantings to establish, they still end up being very diverse.  As our plantings mature, fire and grazing management helps suppress the dominance of big bluestem and indiangrass and maintain high plant diversity.

2.  Weed Control in New Restored Prairies

Weed control strategies for new plantings also vary greatly between Nachusa Grasslands and the Platte River Prairies.  At Nachusa, Bill and his crew walk every inch of new plantings multiple times each year until the native plant community is well established.  They remove (by pulling or spraying) every invasive plant they find – focusing mostly on perennial legumes such as birds foot trefoil, crown vetch, and sweet clover.  Once the native community is established, they can relax a little, but they still watch each site very closely.  In some cases, they’ve not been able to keep up with the pressure from invasive plants and they’ve made the difficult decision to just give up and start over, rather than fighting a losing battle for years.

The soybean field on the left is a former restored prairie that just never established as Bill hoped, so he and his crew made the decision to start over.  After it is farmed for a few years, they'll try again.

The soybean field on the left is a former restored prairie at Nachusa Grasslands that never established as Bill hoped, so he and his crew made the decision to start over. After it is farmed for a few years, they’ll try again.  While it’s a lot of work to start over, it’s less work than many years of fighting weeds and never winning.

Our weed control on the Platte River Prairies looks much different.  We don’t really have problems with perennial legumes or other non-native forbs.  In fact, we pretty much ignore sweet clover, and most other “weeds” during the establishment phase of a new prairie are annuals such as foxtail, marestail, and annual sunflower that just fade away as perennial prairie plants take over.  Our major fears have to do with perennial invasive grasses, such as smooth brome and Kentucky bluegrass, and we deal with those mainly by suppressing them with fire and grazing management.  We also worry about deciduous trees, such as Siberian elms, that we have to control with herbicides because fire and grazing don’t do the job.  However, during the first few years of establishment, while Bill and his crew are painstakingly patrolling their sites, we can mostly ignore our new plantings – except for an occasional prescribed burn to limit the buildup of thatch.

3. Overseeding

A third difference between our sites has to do with overseeding.  In both remnant (never plowed) and restored prairies, we occasionally want to add missing plant species.  At Nachusa, they just burn the prairie and throw the seed out on the bare ground – and it works!  In our drier Platte River Prairies, we’ve not had very much luck with that strategy.  Using grazing to weaken the dominant grasses seems to help us get better establishment of new plants because it suppresses competition for moisture and other resources.  Bill’s crew doesn’t seem to have to worry about that – even in sites with lots of grass.

Bill (left) oversees some "data collection" during this year's Grassland Restoration Network, which was done to help tour participants evaluate an overseeding project.  Becky and Hank Hartman (volunteer stewards) have used repeated overseeding to transform an area from exclusively grasses to one with very nice wildflower diversity.

Bill (left) oversees some “data collection” during this year’s Grassland Restoration Network, which was done to help tour participants evaluate an overseeding project. Becky and Hank Hartman (volunteer stewards) have used repeated overseeding to transform an area from exclusively grasses to one with very nice wildflower diversity.

 

Mike Konen (in orange) from Northern Illinois University talks about soils during this year's workshop at Nachusa.  He is standing in prairie that is going to be grazed by the newly introduced herd of bison.  Grazing will give Bill and I one more thing to compare notes on...

Mike Konen (in orange) from Northern Illinois University talks about soils during this year’s workshop at Nachusa. He is standing in prairie that is going to be grazed by a newly-introduced herd of bison (the bison should knock the height of that indiangrass down some…) Now that Nachusa is using grazing  as a management too, Bill and I will have one more thing to compare notes on…

Bill and I have done extensive experimentation to come up with effective prairie restoration and management strategies at our respective sites, and we continue to adapt as we go along.  We can learn from each others’ experiences, but there is also much that doesn’t translate well between sites.  Some of that is due to the distance between Nachusa and the Platte, and the corresponding differences in climate and soils.  However, even prairies that are much closer together can respond very differently to management and restoration tactics.  Soil types, seed banks, topography, management history, landscape context, and many other factors combine to give every prairie it’s own “personality”.

It might seem overwhelming to learn that every prairie requires a unique set of restoration and management strategies, but it’s really not that bad.  There are a still a lot of commonalities between prairies – just like there are many similarities between Bill and me.  However, just as you would need to consider differences between Bill’s personality and mine in terms if you wanted a positive response from us, the same holds with prairies.  (If you want a favor from Bill, you might want to invite him to a little gathering and serve good beer.  On the flip side, if you want something from me, pizza would make a better bribe, and while I’m not against parties, the less small talk needed, the better…)

Above all, beware of anyone who tells you’ve they’ve figured out the magic formula for how to manage or restore prairies.  It’s just not possible.  Instead, take a look at what others do, learn from their experiences, and then experiment with a variety of techniques at your own site.  It won’t take long to figure out what moves your prairie in the direction you want.   Fortunately, unlike Bill (I’m kidding!) prairies are pretty forgiving, so if you try something and it doesn’t work, they aren’t likely to hold a grudge.

P.S. Bill will be appalled that I’m giving him so much credit for the work at Nachusa.  Clearly, both Bill and I have crews of staff and volunteers that do most of the work and much of the thinking.  For this post, however, I was trying to build an illustration of personalities in people and personalities in prairies.  It was a lot easier to do that by focusing just on Bill and me.  Please understand that the ideas and work of Nelson, Cody, Mardell, Hank, Becky, Karen, Al, Bernie, Jay, Susan, Leah, and many others are represented here as well.  (There, does that make you happier Bill?)